original description
Seninski, K. (1905). Novyya Dannyya o neogenovych' plastach' Yugozapadnogo Zakavkaz'ya. <em>Trudy Obshchestva Estestvoispytateley pri Imperatorskom Yuryevskom Universitete.</em> 16, 1-80.
page(s): 58-59, textfig. 1 [details]
new combination reference
Harzhauser, M.; Mandic, O. (2008). Neogene lake systems of Central and South-Eastern Europe: Faunal diversity, gradients and interrelations. Palaeogeography, Palaeoclimatology, Palaeoecology. 260(3-4): 417-434., available online at https://doi.org/10.1016/j.palaeo.2007.12.013
page(s): database [details]
From editor or global species database
Type locality P'ok'veshi ("Pakweschi"), Abkhazia, Georgia; Duab beds, middle to late Kimmerian, early Pliocene [details]