From editor or global species database
Type locality Beaurains-lès-Noyon ("Beaurein"), Oise, and Épernay ("Épernai"), Marne, France, and London (Charlton) and London (Woolwich) ("Wolwich"), London, and Southfleet, Kent, United Kingdom; late Thanetian to early Ypresian, late Paleocene to early Eocene [details]