basis of recordOlsson, A. A. (1922). The Miocene of northern Costa Rica with notes on its general stratigraphic relations. <em>Bulletins of American Paleontology.</em> 9(39): 173-481 [1-309], pls 1-32 [4-35]., available online athttps://www.biodiversitylibrary.org/page/46040597[details]
Present Inaccurate Introduced: alien Containing type locality
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