From editor or global species database
Type locality Originally as "[dans un terrain de calcaire d'eau douce qui limite] au sud-est le bassin de Toulouse" with no exact locality given, but specified as Mas-Saintes-Puelles, Aude, France (Calcaires de Mas-Saintes-Puelles, Priabonian, late Eocene) by Schnabel (2005). [details]