original description
Roman F. (1912 ["1910"]). Faune saumâtre du Sannoisien du Gard. <em>Bulletin de la Société Géologique de France.</em> ser. 4, 10: 927-955, pls 22-24., available online at https://biodiversitylibrary.org/page/31097480
page(s): 929, pl. 23, figs 6-6c [details]
From editor or global species database
Type locality Beaucaire (Butte Iouton), Gard, France; late Priabonian, late Eocene [details]