original description
Andrusov, N. (1902). Materialy k poznaniyu Prikaspiyskogo neogena. Akchagyl'skiye plasty. <em>Trudy Geologicheskago Komiteta, novaya seriya.</em> 15(4): 1-153.
page(s): 77, 134, pl. 4, fig. 6 [details]
From editor or global species database
Type locality "Inderskiya gory (Belaya Rostosh', r. Ural' u Indera, No. 34), Krasnovodskoye plato (Syuyryumchek')" [Belaya Rostosh, rivers Ural and Inder and exposure No. 34, Inder mountains, W Kazakhstan; Syuyryumchek, Krasnovodsk plateau, Turkmenistan]; Akchagylian [details]