original description
Andrusov, N. (1902). Materialy k poznaniyu Prikaspiyskogo neogena. Akchagyl'skiye plasty. <em>Trudy Geologicheskago Komiteta, novaya seriya.</em> 15(4): 1-153.
page(s): 77-78, 134-135, pl. 4, fig. 8 [details]
From editor or global species database
Type locality "Kuk'-Dzhul' u Krasnovodska" [Kuk'-Dzhul' at Krasnovodsk plateau], Turkmenistan; Akchagylian, late Pliocene-early Pleistocene [details]