original description
Andrusov N. (1890). Kerchenskiy izvestnyak i yego fauna. <em>Zapiski Imperatorskago S.-Petersburgskago Mineralogicheskago Obshchestva, seriya 2.</em> 26: 193-344.
page(s): 246, pl. 2, fig. 4 [details]
From editor or global species database
Type locality "Yuzhnyy bereg' Churubashskago ozera, mezhdu Kapkanami i Novym' Karantinom', Mitridat', Chegene, Ossoviny; [...] Kapkany, [...] Katerles'" [southern coast of Lake Churubash, between Kapkany and Novyy Karantyn, Mt. Mithridat, Chegene, Osovyny, Kapkany and Katerles, Kerch Peninsula, Crimea]; Maeotian, late Miocene [details]