original description
Andrusov, N. (1902). Materialy k poznaniyu Prikaspiyskogo neogena. Akchagyl'skiye plasty. <em>Trudy Geologicheskago Komiteta, novaya seriya.</em> 15(4): 1-153.
page(s): 86-87, 138-139, pl. 4, figs 16-20 [details]
From editor or global species database
Type locality "Akchagyl', Kegnya-Arap', Chir'-yurt' (?), Belaya Rostosh', Inder' No. 38, Brusyanoy Obrag', Utva No. 3 i No. 5" [Akchagyl' and Kegnya-Arap', Turkmenistan; Kizilyurt, Dagestan, Russia (?); Belaya Rostosh', Inder exposure No. 38, Brusyanoy, Utva exposures no. 3 and 5, Kazakhstan] [details]