original description
Andrusov, N. (1917). Ponticheskiy yarus [The Pontian horizon]. Anonymous (Ed.) Geologiya S.S.S.R. Tom IV2, Kaynozoyskaya gruppa. Otdel II (Neogen). Chast' II (Pliotsen). Vyp. 2. Izdaniye Geologicheskogo Komiteta, Leningrad, 1-51 pp.
page(s): pl. 1, fig. 1 [details]
From editor or global species database
Type locality Beslakhuba (at Ghalidzga river), Abkhazia [details]