original description
Papp, A. (1979). Zur Kenntnis neogener Süßwasserfaunen in Attika (Griechenland). Annales Géologiques des Pays Helléniques. 29, 664-678.
page(s): 672, pl. 3, figs 1-10 [details] Available for editors [request]
From editor or global species database
Type locality Milesi ("Milessi"), East Attica, Greece; Tortonian (mammal zone MN 10), late Miocene [details]