original description
Oppenheim, P. (1919). Das Neogen in Kleinasien. <em>Zeitschrift der Deutschen Geologischen Gesellschaft.</em> 70, 1–210, 12 pls.
page(s): 117-119, pl. 5, fig. 2, 2a [details]
From editor or global species database
Type locality Tlos, Mugla, Turkey; Eşen Formation (Kocaçay Member), Zanclean, early Pliocene [details]