original description
Andrusov, N. (1902). Materialy k poznaniyu Prikaspiyskogo neogena. Akchagyl'skiye plasty. <em>Trudy Geologicheskago Komiteta, novaya seriya.</em> 15(4): 1-153.
page(s): 81-84, 136-138, pl. 5, figs 3-6, 30, 31, 33 [details]
From editor or global species database
Type locality Numerous localities in Turkmenistan (Krasnovodsk plateau), Russia (Dagestan, Chechnya regions), Georgia (Eldar steppe) and western Kazakhstan; Akchagylian, late Pliocene-early Pleistocene [details]