original description
Andrusov, N. (1902). Materialy k poznaniyu Prikaspiyskogo neogena. Akchagyl'skiye plasty. <em>Trudy Geologicheskago Komiteta, novaya seriya.</em> 15(4): 1-153.
page(s): 80, 136, pl. 4, fig. 11 [details]
From editor or global species database
Type locality "Kukurt'", Krasnovodsk plateau, Turkmenistan; Akchagylian, late Pliocene-early Pleistocene [details]