original description
Stevanović, P.M. (1951). Pontische Stufe im engeren Sinne - obere Congerienschichten Serbiens und der angrenzenden Gebiete. <em>Sprska Akademija nauka, posebna izdanja.</em> 187(Geološki institut 2): 1-361.
page(s): 212-213, 326-327, pl. 2, fig. 8 [details]
From editor or global species database
Type locality "Nordserbien, Orešac I a. d. Donau, Pontische Stufe s. str., Orešacsande" [Northern Serbia, Orešac I at the Danube, Pontian stage s.str., Orešac sands] [note that in the figure caption the type locality is given as "Brestovnik a. d. Donau"] [details]