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Hershler, R.; Liu, H.-P.; Gustafson, D. L. (2008). A second species of Pyrgulopsis (Hydrobiidae) from the Missouri River basin, with molecular evidence supporting faunal origin through Pliocene stream capture across the northern continental divide. Journal of Molluscan Studies. 74(4): 403-413.
Hershler, R.; Liu, H.-P.; Gustafson, D. L. (2008). A second species of Pyrgulopsis (Hydrobiidae) from the Missouri River basin, with molecular evidence supporting faunal origin through Pliocene stream capture across the northern continental divide. Journal of Molluscan Studies. 74(4): 403-413.