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Harzhauser, M.; Landau, B. M. (2025). The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea—an overwhelming diversity. Zootaxa. 5630(1): 1-142.
508325
urn:lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31 [view]
Harzhauser, M.; Landau, B. M.
2025
The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea—an overwhelming diversity
Zootaxa
5630(1): 1-142
Publication
RIS (EndNote, Reference Manager, ProCite, RefWorks)
BibTex (BibDesk, LaTeX)
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2025-05-01 17:15:36Z
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Acirsa fusiformis Harzhauser & Landau, 2025 † (original description)
Acirsa koskeridouae Harzhauser & Landau, 2025 † (original description)
Acirsa miopedemontana Sacco, 1891 † (basis of record)
Acirsa plana Csepreghy-Meznerics, 1950 † (basis of record)
Acrilloscala austriaca Harzhauser & Landau, 2025 † (original description)
Acrilloscala fortior Harzhauser & Landau, 2025 † (original description)
Acrilloscala propugnaculum Harzhauser & Landau, 2025 † (original description)
Acrilloscala subuloprisca (Sacco, 1891) † (basis of record)
Amaea amoena (R. A. Philippi, 1844) † (new combination reference)
Amaea grunerti Harzhauser & Landau, 2025 † (original description)
Amaea hoernesi (De Gregorio, 1889) † (basis of record)
Amaea scaberrima (Michelotti, 1840) † accepted as Discoscala scaberrima (Michelotti, 1840) † (new combination reference)
Anguloacrilla Harzhauser & Landau, 2025 † (original description)
Anguloacrilla karamanensis Harzhauser & Landau, 2025 † (original description)
Anguloacrilla kimakowiczi (O. Boettger, 1896) † (basis of record)
Anguloacrilla lima Harzhauser & Landau, 2025 † (original description)
Anguloacrilla schwarzhansi Harzhauser & Landau, 2025 † (original description)
Cirsotrema crassicostanomalum (Sacco, 1891) † (basis of record)
Cirsotrema crassicostanomalum (Sacco, 1891) † (subsequent type designation)
Cirsotrema dertonense (Sacco, 1891) † (basis of record)
Cirsotrema dertonense (Sacco, 1891) † (subsequent type designation)
Cirsotrema duciei var. crassicostanomala Sacco, 1891 † accepted as Cirsotrema crassicostanomalum (Sacco, 1891) † (new combination reference)
Cirsotrema fontannesi Cossmann & Peyrot, 1922 † (basis of record)
Cirsotrema hornesi (De Gregorio, 1889) † (basis of record)
Cirsotrema kokayi Kovács, Leél-Őssy & Vicián, 2023 † (additional source)
Cirsotrema loerentheyi (O. Boettger, 1902) † (basis of record)
Cirsotrema menkveldae Pfister & Wegmüller, 2007 † (basis of record)
Cirsotrema microperforata Harzhauser & Landau, 2025 † (original description)
Cirsotrema nudum Harzhauser & Landau, 2025 † (original description)
Cirsotrema pollerspoecki Harzhauser & Landau, 2025 † (original description)
Cirsotrema pumiceum (Brocchi, 1814) (additional source)
Cirsotrema schneideri Harzhauser & Landau, 2025 † (original description)
Clathrus corrugatus Harzhauser & Landau, 2025 † (original description)
Clathrus detractus Cossmann, 1912 † accepted as Nitidiscala detracta (Cossmann, 1912) † (new combination reference)
Claviscala norica Harzhauser & Landau, 2014 † (additional source)
Cylindriscala acus (R. B. Watson, 1883) (additional source)
Cylindriscala amandae (O. Boettger, 1906) † (basis of record)
Cylindriscala bellissima Harzhauser & Landau, 2025 † (original description)
Cylindriscala rugata Harzhauser & Landau, 2025 † (original description)
Discoscala Sacco, 1890 (basis of record)
Funiscala infans (O. Boettger, 1906) † (basis of record)
Gregorioiscala ascialis Harzhauser & Landau, 2025 † (original description)
Hemiacirsa de Boury, 1890 (redescription)
Hemiacirsa abiaeformis Harzhauser & Landau, 2025 † (original description)
Hemiacirsa drevermanni (Cossmann, 1912) † (basis of record)
Hemiacirsa incusa Harzhauser & Landau, 2025 † (original description)
Hemiacirsa miocenica (Seguenza, 1880) † (basis of record)
Hemiacirsa oscari Cossmann, 1912 † (basis of record)
Hemiacirsa superelongata Harzhauser & Landau, 2025 † (original description)
Hirtoscala ceruchus Harzhauser & Landau, 2025 † (original description)
Hirtoscala cornucopiae Harzhauser & Landau, 2025 † (original description)
Hirtoscala diplommatinaformis Harzhauser & Landau, 2025 † (original description)
Hirtoscala hastata Harzhauser & Landau, 2025 † (original description)
Hirtoscala holcovae Harzhauser & Landau, 2025 † (original description)
Hirtoscala miofrondiculoides (Sacco, 1891) † (basis of record)
Hirtoscala pharos Harzhauser & Landau, 2025 † (original description)
Hirtoscala steiningeri Harzhauser & Landau, 2025 † (original description)
Hirtoscala uncinata Harzhauser & Landau, 2025 † (original description)
Hyaloscala occidentalis Harzhauser & Landau, 2025 † (original description)
Hyaloscala rainesi Harzhauser & Landau, 2025 † (original description)
Hyaloscala rustica Harzhauser & Landau, 2025 † (original description)
Librariscala Kilburn, 1985 (taxonomy source)
Librariscala costatissima Harzhauser & Landau, 2025 † (original description)
Mazescala alata Harzhauser & Landau, 2025 † (original description)
Mazescala kostejana (O. Boettger, 1902) † (basis of record)
Nitidiscala baluki Harzhauser & Landau, 2025 † (original description)
Nitidiscala bonfittoi Harzhauser & Landau, 2025 † (original description)
Nitidiscala browni Harzhauser & Landau, 2025 † (original description)
Nitidiscala connexicosta Harzhauser & Landau, 2025 † (original description)
Nitidiscala detracta (Cossmann, 1912) † (basis of record)
Nitidiscala oligocostata (Bałuk, 1975) † (basis of record)
Nitidiscala pultoastensis (Sacco, 1891) † (basis of record)
Nodiscala bimonilifera (O. Boettger, 1902) † accepted as Nodiscala scacchii (M. Hörnes, 1856) † (source of synonymy)
Nodiscala macilenta (Cossmann, 1912) † (basis of record)
Nodiscala microscopica (O. Boettger, 1906) † (basis of record)
Nodiscala rugatina de Boury, 1912 † accepted as Nodiscala scacchii (M. Hörnes, 1856) † (source of synonymy)
Nodiscala scacchii (M. Hörnes, 1856) † (basis of record)
Nodiscala scacchii (M. Hörnes, 1856) † (subsequent type designation)
Nodiscala transsylvanica (O. Boettger, 1902) † (basis of record)
Opaliopsis compacta Harzhauser & Landau, 2025 † (original description)
Papuliscala aquaensis Harzhauser & Landau, 2025 † (original description)
Papuliscala hoffmani Harzhauser & Landau, 2025 † (original description)
Papuliscala maysi Harzhauser & Landau, 2025 † (original description)
Papuliscala parvicancellata Harzhauser & Landau, 2025 † (original description)
Papuliscala plioligustica (Sacco, 1891) † (new combination reference)
Papuliscala praeelongata Harzhauser & Landau, 2025 † (original description)
Papuliscala subsoluta (O. Boettger, 1907) † (new combination reference)
Parviscala abdita Harzhauser & Landau, 2025 † (original description)
Parviscala badensis Harzhauser & Landau, 2025 † (original description)
Parviscala lacinia Harzhauser & Landau, 2025 † (original description)
Parviscala paratethyca Harzhauser & Landau, 2025 † (original description)
Pliciscala (Funiscala) microscopica O. Boettger, 1906 † accepted as Nodiscala microscopica (O. Boettger, 1906) † (basis of record)
Pliciscala (Turriscala) torulosa var. acutinodosa Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. apuniculata Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. atava Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. cingulata Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. convexiuscula Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. perconica Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Pliciscala (Turriscala) torulosa var. rovasendae Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (new combination reference)
Pliciscala (Turriscala) torulosa var. supervaricosa Sacco, 1891 † accepted as Turriscala torulosa (Brocchi, 1814) † (source of synonymy)
Austria for Acirsa koskeridouae Harzhauser & Landau, 2025 † 
Austria for Acrilloscala austriaca Harzhauser & Landau, 2025 † 
Austria for Acrilloscala fortior Harzhauser & Landau, 2025 † 
Austria for Acrilloscala propugnaculum Harzhauser & Landau, 2025 † 
Austria for Amaea grunerti Harzhauser & Landau, 2025 † 
Austria for Amaea hoernesi (De Gregorio, 1889) † 
Austria for Anguloacrilla schwarzhansi Harzhauser & Landau, 2025 † 
Austria for Cirsotrema microperforata Harzhauser & Landau, 2025 † 
Austria for Cirsotrema nudum Harzhauser & Landau, 2025 † 
Austria for Cirsotrema schneideri Harzhauser & Landau, 2025 † 
Austria for Clathrus corrugatus Harzhauser & Landau, 2025 † 
Austria for Cylindriscala bellissima Harzhauser & Landau, 2025 † 
Austria for Hemiacirsa abiaeformis Harzhauser & Landau, 2025 † 
Austria for Hemiacirsa incusa Harzhauser & Landau, 2025 † 
Austria for Hirtoscala ceruchus Harzhauser & Landau, 2025 † 
Austria for Hirtoscala hastata Harzhauser & Landau, 2025 † 
Austria for Hirtoscala holcovae Harzhauser & Landau, 2025 † 
Austria for Hirtoscala steiningeri Harzhauser & Landau, 2025 † 
Austria for Hirtoscala uncinata Harzhauser & Landau, 2025 † 
Austria for Hyaloscala occidentalis Harzhauser & Landau, 2025 † 
Austria for Hyaloscala rustica Harzhauser & Landau, 2025 † 
Austria for Librariscala costatissima Harzhauser & Landau, 2025 † 
Austria for Mazescala alata Harzhauser & Landau, 2025 † 
Austria for Papuliscala aquaensis Harzhauser & Landau, 2025 † 
Austria for Papuliscala parvicancellata Harzhauser & Landau, 2025 † 
Austria for Parviscala abdita Harzhauser & Landau, 2025 † 
Austria for Parviscala badensis Harzhauser & Landau, 2025 † 
Austria for Parviscala paratethyca Harzhauser & Landau, 2025 † 
Austria for Scalaria (Hirtoscala) miofrondiculoides Sacco, 1891 † 
Austria for Scalaria scacchii M. Hörnes, 1856 † 
Austria for Subuliscala breitenbergeri Harzhauser & Landau, 2025 † 
Austria for Subuliscala pseudobanoni Harzhauser & Landau, 2025 † 
Czech Republic for Cylindriscala rugata Harzhauser & Landau, 2025 † 
Czech Republic for Gregorioiscala ascialis Harzhauser & Landau, 2025 † 
Czech Republic for Hirtoscala diplommatinaformis Harzhauser & Landau, 2025 † 
Czech Republic for Nitidiscala connexicosta Harzhauser & Landau, 2025 † 
Czech Republic for Opaliopsis compacta Harzhauser & Landau, 2025 † 
Czech Republic for Papuliscala hoffmani Harzhauser & Landau, 2025 † 
Czech Republic for Papuliscala maysi Harzhauser & Landau, 2025 † 
Czech Republic for Papuliscala praeelongata Harzhauser & Landau, 2025 † 
Czech Republic for Punctiscala moravica Harzhauser & Landau, 2025 † 
France for Cirsotrema fontannesi Cossmann & Peyrot, 1922 † 
Germany for Cirsotrema pollerspoecki Harzhauser & Landau, 2025 † 
Germany for Sthenorytis simoni Harzhauser & Landau, 2025 † 
Italy for Cirsotrema duciei var. crassicostanomala Sacco, 1891 † 
Italy for Scalaria (Cirsotrema) pumiceum var. ingauna Sacco, 1891 † 
Italy for Scalaria (Clathrus) communis var. pultoastensis Sacco, 1891 † 
Poland for Hirtoscala cornucopiae Harzhauser & Landau, 2025 † 
Poland for Scala polonica Harzhauser & Landau, 2025 † 
Romania for Acirsa fusiformis Harzhauser & Landau, 2025 † 
Romania for Anguloacrilla lima Harzhauser & Landau, 2025 † 
Romania for Hemiacirsa oscari Cossmann, 1912 † 
Romania for Hemiacirsa superelongata Harzhauser & Landau, 2025 † 
Romania for Hirtoscala pharos Harzhauser & Landau, 2025 † 
Romania for Nitidiscala baluki Harzhauser & Landau, 2025 † 
Romania for Nitidiscala bonfittoi Harzhauser & Landau, 2025 † 
Romania for Nitidiscala browni Harzhauser & Landau, 2025 † 
Romania for Parviscala lacinia Harzhauser & Landau, 2025 † 
Switzerland for Cirsotrema menkveldae Pfister & Wegmüller, 2007 † 
Turkey for Anguloacrilla karamanensis Harzhauser & Landau, 2025 † 










































































































































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