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Korshunova, T.; Fletcher, K.; Martynov, A. (2025). The endless forms are the most differentiated — how taxonomic pseudo-optimization masked natural diversity and evolution: the nudibranch case. Zoological Journal of the Linnean Society. 204(4): zlaf057: 1-93.
514054
10.1093/zoolinnean/zlaf057 [view]
Korshunova, T.; Fletcher, K.; Martynov, A.
2025
The endless forms are the most differentiated — how taxonomic pseudo-optimization masked natural diversity and evolution: the nudibranch case
Zoological Journal of the Linnean Society
204(4): zlaf057: 1-93
Publication
Stated date: 18 August 2025
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In this study, a global-encompassing, fine-scale differentiated taxonomy is consistently presented from the high level of a major nudibranch suborder, Aeolidacea, and then consequentially to lower levels of superfamilies, families, genera, and species to conjoin the underlying evolutionary pathways of aeolidacean nudibranchs and systematic representation to a maximal degree. The suborder Aeolidacea is reinstated and the superfamily system of Aeolidacea is reformed; 10 aeolidacean superfamilies are recognized encompassing 29 families in an all-family Synopsis, and the superfamily Embletonioidea is included in addition. Two new families, Chudidae fam. nov. and Hantazuidae fam. nov., are described, which show unique morphological and molecular patterns. Seven new species are described within various superfamilies, and five species are reinstated. The genus-level taxonomy of the family Coryphellidae is revisited, 11 genera are recognized, including nine reinstated and two new genera. We show that recent synonymization of all valid Coryphellidae genera into just a single genus was based on an erroneous phylogenetic analysis, the careless treatment of morphological characters, and established preconceptions. Uniting fine-scale morphological and molecular data, order level nudibranch taxonomy is refined, order Nudibranchia is restricted, and order Doridida is reinstated. Within the order Nudibranchia, five suborders are recognized: Arminacea, restricted, reinstated; Tritoniacea, reinstated; Dendronotacea, restricted, reinstated; Janolacea, reinstated; and Aeolidacea, revised, reinstated.
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Abronicidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Aeolidina (status source)
Amphorina Quatrefages, 1844 (status source)
Amphorina amazighi (Tamsouri, Carmona, Moukrim & Cervera, 2015) (new combination reference)
Amphorina andra Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Amphorina farrani (Alder & Hancock, 1844) (additional source)
Amphorina linensis (García-Gómez, Cervera & F. J. Garcia, 1990) (additional source)
Amphorina pallida (Alder & Hancock, 1842) (additional source)
Amphorina viriola Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Annulorhina K. P. Rao, 1968 (status source)
Annulorhina mandapamensis K. P. Rao, 1968 (status source)
Apataoidea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (basis of record)
Arminina (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Borealea nobilis (A. E. Verrill, 1880) (status source)
Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Capellinia Trinchese, 1873 (basis of record)
Capellinia vittata (Alder & Hancock, 1842) (new combination reference)
Catriona aurantia (Alder & Hancock, 1842) (status source)
Chudidae Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo humanistica Korshunova, Fletcher & Martynov, 2025 (original description)
Chudoidea Korshunova, Fletcher & Martynov, 2025 (original description)
Corrupta Korshunova, Fletcher & Martynov, 2025 (original description)
Coryphella alexanderi Ekimova, 2022 accepted as Corrupta alexanderi (Ekimova, 2022) (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) accepted as Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 accepted as Occidenthella athadona (Bergh, 1875) (status source)
Coryphella borealis Odhner, 1922 accepted as Gulenia borealis (Odhner, 1922) (status source)
Coryphella browni Picton, 1980 accepted as Fjordia browni (Picton, 1980) (status source)
Coryphella californica Bergh, 1904 accepted as Flabellinopsis iodinea (J. G. Cooper, 1863) (status source)
Coryphella chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 accepted as Orienthella cooperi (T. D. A. Cockerell, 1901) (status source)
Coryphella falklandica Eliot, 1907 accepted as Itaxia falklandica (Eliot, 1907) (status source)
Coryphella fogata (Millen & Hermosillo, 2007) accepted as Orienthella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) accepted as Fjordia insolita (García-Gómez & Cervera, 1989) (status source)
Coryphella lineata (Lovén, 1846) accepted as Fjordia lineata (Lovén, 1846) (status source)
Coryphella longicaudata O'Donoghue, 1922 (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella nobilis A. E. Verrill, 1880 accepted as Borealea nobilis (A. E. Verrill, 1880) (status source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella trilineata O'Donoghue, 1921 accepted as Orienthella trilineata (O'Donoghue, 1921) (status source)
Coryphella trophina (Bergh, 1890) accepted as Himatina trophina (Bergh, 1890) (status source)
Coryphella verta Ev. Marcus, 1970 accepted as Orienthella verta (Marcus, Ev. 1970) (status source)
Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025 (original description)
Dendronotina (status source)
Diaphoreolis flavovulta (MacFarland, 1966) (status source)
Diaphoreolis lagunae (O'Donoghue, 1926) (status source)
Diaphoreolis stipata (Alder & Hancock, 1843) (status source)
Diaphoreolis zvezda Korshunova, Fletcher, Bakken & Martynov, 2023 (status source)
Doridida (status source)
Dunga Eliot, 1902 (status source)
Embletonioidea Pruvot-Fol, 1954 (basis of record)
Eubranchidae Odhner, 1934 (additional source)
Eubranchopsis virginalis Baba, 1949 (status source)
Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchulus olivaceus (O'Donoghue, 1922) (status source)
Eubranchulus rupium (Møller, 1842) (status source)
Eubranchus novik Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024 accepted as Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchus rubropunctatus Edmunds, 1969 accepted as Produnga rubropunctata (Edmunds, 1969) (status source)
Fionoidea J. E. Gray, 1857 (basis of record)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Fjordia browni (Picton, 1980) (status source)
Fjordia capensis (Thiele, 1925) (status source)
Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Fjordia insolita (García-Gómez & Cervera, 1989) (status source)
Fjordia lineata (Lovén, 1846) (status source)
Galvinella Eliot, 1907 (status source)
Galvinella adarensis (Odhner, 1934) (basis of record)
Galvinella antarctica Eliot, 1907 (status source)
Galvinella glacialis Thiele, 1912 (status source)
Goniaeolididae Odhner, 1907 (status source)
Gulenia borealis (Odhner, 1922) (status source)
Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Hantazuia Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuidae Korshunova, Fletcher & Martynov, 2025 (original description)
Himatina Thiele, 1931 (status source)
Itaxia falklandica (Eliot, 1907) (status source)
Janolina (status source)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Microchlamylla gracilis (Alder & Hancock, 1844) (status source)
Notaeolidioidea Eliot, 1910 (basis of record)
Nudibranchus Martynov, 1998 (status source)
Nudibranchus exiguus (Alder & Hancock, 1848) (basis of record)
Occidenthella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Occidenthella athadona (Bergh, 1875) (status source)
Orienthella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Orienthella cooperi (T. D. A. Cockerell, 1901) (status source)
Orienthella fogata (Millen & Hermosillo, 2007) (status source)
Orienthella piunca (Marcus, Er., 1961) (status source)
Orienthella trilineata (O'Donoghue, 1921) (status source)
Phestilla Bergh, 1874 (status source)
Phestilla arnoldi Mehrotra & Caballer, 2024 (status source)
Aeolidina (status source)
Amphorina Quatrefages, 1844 (status source)
Amphorina amazighi (Tamsouri, Carmona, Moukrim & Cervera, 2015) (new combination reference)
Amphorina andra Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Amphorina farrani (Alder & Hancock, 1844) (additional source)
Amphorina linensis (García-Gómez, Cervera & F. J. Garcia, 1990) (additional source)
Amphorina pallida (Alder & Hancock, 1842) (additional source)
Amphorina viriola Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Annulorhina K. P. Rao, 1968 (status source)
Annulorhina mandapamensis K. P. Rao, 1968 (status source)
Apataoidea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (basis of record)
Arminina (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Borealea nobilis (A. E. Verrill, 1880) (status source)
Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Capellinia Trinchese, 1873 (basis of record)
Capellinia vittata (Alder & Hancock, 1842) (new combination reference)
Catriona aurantia (Alder & Hancock, 1842) (status source)
Chudidae Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo humanistica Korshunova, Fletcher & Martynov, 2025 (original description)
Chudoidea Korshunova, Fletcher & Martynov, 2025 (original description)
Corrupta Korshunova, Fletcher & Martynov, 2025 (original description)
Coryphella alexanderi Ekimova, 2022 accepted as Corrupta alexanderi (Ekimova, 2022) (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) accepted as Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 accepted as Occidenthella athadona (Bergh, 1875) (status source)
Coryphella borealis Odhner, 1922 accepted as Gulenia borealis (Odhner, 1922) (status source)
Coryphella browni Picton, 1980 accepted as Fjordia browni (Picton, 1980) (status source)
Coryphella californica Bergh, 1904 accepted as Flabellinopsis iodinea (J. G. Cooper, 1863) (status source)
Coryphella chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 accepted as Orienthella cooperi (T. D. A. Cockerell, 1901) (status source)
Coryphella falklandica Eliot, 1907 accepted as Itaxia falklandica (Eliot, 1907) (status source)
Coryphella fogata (Millen & Hermosillo, 2007) accepted as Orienthella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) accepted as Fjordia insolita (García-Gómez & Cervera, 1989) (status source)
Coryphella lineata (Lovén, 1846) accepted as Fjordia lineata (Lovén, 1846) (status source)
Coryphella longicaudata O'Donoghue, 1922 (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella nobilis A. E. Verrill, 1880 accepted as Borealea nobilis (A. E. Verrill, 1880) (status source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) accepted as Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphella trilineata O'Donoghue, 1921 accepted as Orienthella trilineata (O'Donoghue, 1921) (status source)
Coryphella trophina (Bergh, 1890) accepted as Himatina trophina (Bergh, 1890) (status source)
Coryphella verta Ev. Marcus, 1970 accepted as Orienthella verta (Marcus, Ev. 1970) (status source)
Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025 (original description)
Dendronotina (status source)
Diaphoreolis flavovulta (MacFarland, 1966) (status source)
Diaphoreolis lagunae (O'Donoghue, 1926) (status source)
Diaphoreolis stipata (Alder & Hancock, 1843) (status source)
Diaphoreolis zvezda Korshunova, Fletcher, Bakken & Martynov, 2023 (status source)
Doridida (status source)
Dunga Eliot, 1902 (status source)
Embletonioidea Pruvot-Fol, 1954 (basis of record)
Eubranchidae Odhner, 1934 (additional source)
Eubranchopsis virginalis Baba, 1949 (status source)
Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchulus olivaceus (O'Donoghue, 1922) (status source)
Eubranchulus rupium (Møller, 1842) (status source)
Eubranchus novik Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024 accepted as Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchus rubropunctatus Edmunds, 1969 accepted as Produnga rubropunctata (Edmunds, 1969) (status source)
Fionoidea J. E. Gray, 1857 (basis of record)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Fjordia browni (Picton, 1980) (status source)
Fjordia capensis (Thiele, 1925) (status source)
Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Fjordia insolita (García-Gómez & Cervera, 1989) (status source)
Fjordia lineata (Lovén, 1846) (status source)
Galvinella Eliot, 1907 (status source)
Galvinella adarensis (Odhner, 1934) (basis of record)
Galvinella antarctica Eliot, 1907 (status source)
Galvinella glacialis Thiele, 1912 (status source)
Goniaeolididae Odhner, 1907 (status source)
Gulenia borealis (Odhner, 1922) (status source)
Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Hantazuia Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuidae Korshunova, Fletcher & Martynov, 2025 (original description)
Himatina Thiele, 1931 (status source)
Itaxia falklandica (Eliot, 1907) (status source)
Janolina (status source)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Microchlamylla gracilis (Alder & Hancock, 1844) (status source)
Notaeolidioidea Eliot, 1910 (basis of record)
Nudibranchus Martynov, 1998 (status source)
Nudibranchus exiguus (Alder & Hancock, 1848) (basis of record)
Occidenthella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Occidenthella athadona (Bergh, 1875) (status source)
Orienthella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Orienthella cooperi (T. D. A. Cockerell, 1901) (status source)
Orienthella fogata (Millen & Hermosillo, 2007) (status source)
Orienthella piunca (Marcus, Er., 1961) (status source)
Orienthella trilineata (O'Donoghue, 1921) (status source)
Phestilla Bergh, 1874 (status source)
Phestilla arnoldi Mehrotra & Caballer, 2024 (status source)
Gulf of Cadiz for Piseinotecus soussi Tamsouri, Carmona, Moukrim & Cervera, 2014
Italian part of the Adriatic Sea for Piseinotecus soussi Tamsouri, Carmona, Moukrim & Cervera, 2014
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Okinawa Island for Unidentia kiku Korshunova, Fletcher & Martynov, 2025
Italian part of the Adriatic Sea for Piseinotecus soussi Tamsouri, Carmona, Moukrim & Cervera, 2014
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Okinawa Island for Unidentia kiku Korshunova, Fletcher & Martynov, 2025
Holotype KM1044, geounit Japan, identified as Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1068, geounit Japan, identified as Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Holotype KM883, geounit Southern Ocean, identified as Chudo humanistica Korshunova, Fletcher & Martynov, 2025
Holotype KM954, verbatimGeounit USA, Washington Stat..., identified as Portorchardia candela Korshunova, Fletcher & Martynov, 2025
Holotype ZMMU Op-132.1, geounit Vietnam, identified as Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1068, geounit Japan, identified as Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Holotype KM883, geounit Southern Ocean, identified as Chudo humanistica Korshunova, Fletcher & Martynov, 2025
Holotype KM954, verbatimGeounit USA, Washington Stat..., identified as Portorchardia candela Korshunova, Fletcher & Martynov, 2025
Holotype ZMMU Op-132.1, geounit Vietnam, identified as Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025
