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Richling, I. (2004). Classification of the Helicinidae: Review of morphological characteristics based on a revision of the Costa Rican species and application to the arrangement of the Central American mainland taxa (Mollusca: Gastropoda: Neritopsina). Malacologia. 45(2): 195-440.
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Richling, I.
2004
Classification of the Helicinidae: Review of morphological characteristics based on a revision of the Costa Rican species and application to the arrangement of the Central American mainland taxa (Mollusca: Gastropoda: Neritopsina)
Malacologia
45(2): 195-440
Publication
Date
action
by
Alcadia (Microalcadia) Richling, 2004 represented as Alcadia J. E. Gray, 1840 (original description)
Alcadia (Microalcadia) boeckeleri (Richling, 2001) represented as Alcadia boeckeleri (Richling, 2001) (basis of record)
Alcadia (Microalcadia) hojarasca (Richling, 2001) represented as Alcadia hojarasca (Richling, 2001) (basis of record)
Alcadia (Palliata) A. J. Wagner, 1907 represented as Alcadia J. E. Gray, 1840 (additional source)
Alcadia jamaicensis (G. B. Sowerby II, 1842) (status source)
Ceochasma phrixina F. G. Thompson, 1968 (additional source)
Helicina (Analcadia) A. J. Wagner, 1908 represented as Helicina Lamarck, 1799 (basis of record)
Helicina (Analcadia) dysoni L. Pfeiffer, 1849 represented as Helicina dysoni L. Pfeiffer, 1849 (basis of record)
Helicina (Gemma) beatrix Angas, 1879 represented as Helicina beatrix Angas, 1879 (basis of record)
Helicina (Gemma) beatrix beatrix Angas, 1879 represented as Helicina beatrix beatrix Angas, 1879 (basis of record)
Helicina (Gemma) beatrix riopejensis Richling, 2004 represented as Helicina beatrix riopejensis Richling, 2004 (original description)
Helicina (Gemma) chiquitica (Richling, 2001) represented as Helicina chiquitica (Richling, 2001) (basis of record)
Helicina (Gemma) escondida Richling, 2004 represented as Helicina escondida Richling, 2004 (original description)
Helicina (Gemma) gemma Preston, 1903 represented as Helicina gemma Preston, 1903 (basis of record)
Helicina (Gemma) monteverdensis Richling, 2004 represented as Helicina monteverdensis Richling, 2004 (original description)
Helicina (Gemma) talamancensis (Richling, 2001) represented as Helicina talamancensis (Richling, 2001) (basis of record)
Helicina (Retorquata) funcki costaricensis A. J. Wagner, 1905 accepted as Helicina funcki L. Pfeiffer, 1849 (source of synonymy)
Helicina (Tristramia) echandiensis Richling, 2004 represented as Helicina echandiensis Richling, 2004 (original description)
Helicina (Tristramia) funcki L. Pfeiffer, 1849 represented as Helicina funcki L. Pfeiffer, 1849 (basis of record)
Helicina (Tristramia) pitalensis A. J. Wagner, 1910 represented as Helicina pitalensis A. J. Wagner, 1910 (basis of record)
Helicina (Tristramia) punctisulcata E. von Martens, 1890 represented as Helicina punctisulcata E. von Martens, 1890 (basis of record)
Helicina (Tristramia) punctisulcata cuericiensis Richling, 2004 represented as Helicina punctisulcata cuericiensis Richling, 2004 (original description)
Helicina beatrix confusa (A. J. Wagner, 1908) (basis of record)
Helicina beatrix nicaraguae (A. J. Wagner, 1908) (basis of record)
Helicina chiapensis L. Pfeiffer, 1856 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina chryseis Tristram, 1861 accepted as Pyrgodomus microdinus (Morelet, 1851) (source of synonymy)
Helicina diaphana L. Pfeiffer, 1852 (basis of record)
Helicina elata Shuttleworth, 1852 (basis of record)
Helicina fragilis Morelet, 1851 (basis of record)
Helicina funcki parvidens Pilsbry, 1920 accepted as Helicina funcki L. Pfeiffer, 1849 (source of synonymy)
Helicina lamellosa Guppy, 1867 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina merdigera L. Pfeiffer, 1855 (basis of record)
Helicina mohriana L. Pfeiffer, 1861 (basis of record)
Helicina oweniana L. Pfeiffer, 1849 (basis of record)
Helicina oweniana anozona E. von Martens, 1875 (basis of record)
Helicina oweniana coccinostoma Morelet, 1849 (basis of record)
Helicina rusticella Morelet, 1849 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina semistriata G. B. Sowerby II, 1866 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina tenuis pittieri A. J. Wagner, 1910 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina terryae Rehder, 1940 (basis of record)
Helicina unidentata L. Pfeiffer, 1849 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina vernalis Morelet, 1849 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina vernalis verapazensis A. J. Wagner, 1905 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Lucidella lirata (L. Pfeiffer, 1847) (basis of record)
Pyrgodomus microdinus (Morelet, 1851) (basis of record)
Alcadia (Microalcadia) boeckeleri (Richling, 2001) represented as Alcadia boeckeleri (Richling, 2001) (basis of record)
Alcadia (Microalcadia) hojarasca (Richling, 2001) represented as Alcadia hojarasca (Richling, 2001) (basis of record)
Alcadia (Palliata) A. J. Wagner, 1907 represented as Alcadia J. E. Gray, 1840 (additional source)
Alcadia jamaicensis (G. B. Sowerby II, 1842) (status source)
Ceochasma phrixina F. G. Thompson, 1968 (additional source)
Helicina (Analcadia) A. J. Wagner, 1908 represented as Helicina Lamarck, 1799 (basis of record)
Helicina (Analcadia) dysoni L. Pfeiffer, 1849 represented as Helicina dysoni L. Pfeiffer, 1849 (basis of record)
Helicina (Gemma) beatrix Angas, 1879 represented as Helicina beatrix Angas, 1879 (basis of record)
Helicina (Gemma) beatrix beatrix Angas, 1879 represented as Helicina beatrix beatrix Angas, 1879 (basis of record)
Helicina (Gemma) beatrix riopejensis Richling, 2004 represented as Helicina beatrix riopejensis Richling, 2004 (original description)
Helicina (Gemma) chiquitica (Richling, 2001) represented as Helicina chiquitica (Richling, 2001) (basis of record)
Helicina (Gemma) escondida Richling, 2004 represented as Helicina escondida Richling, 2004 (original description)
Helicina (Gemma) gemma Preston, 1903 represented as Helicina gemma Preston, 1903 (basis of record)
Helicina (Gemma) monteverdensis Richling, 2004 represented as Helicina monteverdensis Richling, 2004 (original description)
Helicina (Gemma) talamancensis (Richling, 2001) represented as Helicina talamancensis (Richling, 2001) (basis of record)
Helicina (Retorquata) funcki costaricensis A. J. Wagner, 1905 accepted as Helicina funcki L. Pfeiffer, 1849 (source of synonymy)
Helicina (Tristramia) echandiensis Richling, 2004 represented as Helicina echandiensis Richling, 2004 (original description)
Helicina (Tristramia) funcki L. Pfeiffer, 1849 represented as Helicina funcki L. Pfeiffer, 1849 (basis of record)
Helicina (Tristramia) pitalensis A. J. Wagner, 1910 represented as Helicina pitalensis A. J. Wagner, 1910 (basis of record)
Helicina (Tristramia) punctisulcata E. von Martens, 1890 represented as Helicina punctisulcata E. von Martens, 1890 (basis of record)
Helicina (Tristramia) punctisulcata cuericiensis Richling, 2004 represented as Helicina punctisulcata cuericiensis Richling, 2004 (original description)
Helicina beatrix confusa (A. J. Wagner, 1908) (basis of record)
Helicina beatrix nicaraguae (A. J. Wagner, 1908) (basis of record)
Helicina chiapensis L. Pfeiffer, 1856 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina chryseis Tristram, 1861 accepted as Pyrgodomus microdinus (Morelet, 1851) (source of synonymy)
Helicina diaphana L. Pfeiffer, 1852 (basis of record)
Helicina elata Shuttleworth, 1852 (basis of record)
Helicina fragilis Morelet, 1851 (basis of record)
Helicina funcki parvidens Pilsbry, 1920 accepted as Helicina funcki L. Pfeiffer, 1849 (source of synonymy)
Helicina lamellosa Guppy, 1867 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina merdigera L. Pfeiffer, 1855 (basis of record)
Helicina mohriana L. Pfeiffer, 1861 (basis of record)
Helicina oweniana L. Pfeiffer, 1849 (basis of record)
Helicina oweniana anozona E. von Martens, 1875 (basis of record)
Helicina oweniana coccinostoma Morelet, 1849 (basis of record)
Helicina rusticella Morelet, 1849 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina semistriata G. B. Sowerby II, 1866 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina tenuis pittieri A. J. Wagner, 1910 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina terryae Rehder, 1940 (basis of record)
Helicina unidentata L. Pfeiffer, 1849 accepted as Lucidella lirata (L. Pfeiffer, 1847) (source of synonymy)
Helicina vernalis Morelet, 1849 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Helicina vernalis verapazensis A. J. Wagner, 1905 accepted as Helicina tenuis L. Pfeiffer, 1849 (source of synonymy)
Lucidella lirata (L. Pfeiffer, 1847) (basis of record)
Pyrgodomus microdinus (Morelet, 1851) (basis of record)
Chiapas for Helicina chiapensis L. Pfeiffer, 1856
Costa Rica for Helicina (Gemma) escondida Richling, 2004
Costa Rica for Helicina (Gemma) monteverdensis Richling, 2004
Costa Rica for Helicina beatrix Angas, 1879
Costa Rica for Helicina funcki L. Pfeiffer, 1849
Costa Rica for Helicina gemma Preston, 1903
Guatemala for Helicina anozona E. von Martens, 1875
Guatemala for Helicina coccinostoma Morelet, 1849
Guatemala for Helicina fragilis Morelet, 1851
Guatemala for Helicina vernalis Morelet, 1849
Guatemala for Helicina vernalis verapazensis A. J. Wagner, 1905
Honduras for Helicina diaphana L. Pfeiffer, 1852
Honduras for Helicina unidentata L. Pfeiffer, 1849
Limon for Alcadia (Leialcadia) beatrix confusa A. J. Wagner, 1908
Limon for Helicina (Gemma) beatrix riopejensis Richling, 2004
Limon for Helicina beatrix riopejensis Richling, 2004
Limon for Oligyra chiquitica Richling, 2001
Nicaragua for Alcadia (Leialcadia) beatrix nicaraguae A. J. Wagner, 1908
Nicaragua for Helicina funcki L. Pfeiffer, 1849
Panama for Helicina funcki L. Pfeiffer, 1849
Puntarenas for Helicina pitalensis A. J. Wagner, 1910
Puntarenas for Helicina tenuis pittieri A. J. Wagner, 1910
San José for Helicina (Tristramia) punctisulcata cuericiensis Richling, 2004
Veracruz for Helicina elata Shuttleworth, 1852
Veracruz for Helicina merdigera L. Pfeiffer, 1855
Veraguas for Helicina funcki L. Pfeiffer, 1849
Yucatan for Helicina lirata L. Pfeiffer, 1847
Yucatan for Helicina tenuis L. Pfeiffer, 1849
Costa Rica for Helicina (Gemma) escondida Richling, 2004
Costa Rica for Helicina (Gemma) monteverdensis Richling, 2004
Costa Rica for Helicina beatrix Angas, 1879
Costa Rica for Helicina funcki L. Pfeiffer, 1849
Costa Rica for Helicina gemma Preston, 1903
Guatemala for Helicina anozona E. von Martens, 1875
Guatemala for Helicina coccinostoma Morelet, 1849
Guatemala for Helicina fragilis Morelet, 1851
Guatemala for Helicina vernalis Morelet, 1849
Guatemala for Helicina vernalis verapazensis A. J. Wagner, 1905
Honduras for Helicina diaphana L. Pfeiffer, 1852
Honduras for Helicina unidentata L. Pfeiffer, 1849
Limon for Alcadia (Leialcadia) beatrix confusa A. J. Wagner, 1908
Limon for Helicina (Gemma) beatrix riopejensis Richling, 2004
Limon for Helicina beatrix riopejensis Richling, 2004
Limon for Oligyra chiquitica Richling, 2001
Nicaragua for Alcadia (Leialcadia) beatrix nicaraguae A. J. Wagner, 1908
Nicaragua for Helicina funcki L. Pfeiffer, 1849
Panama for Helicina funcki L. Pfeiffer, 1849
Puntarenas for Helicina pitalensis A. J. Wagner, 1910
Puntarenas for Helicina tenuis pittieri A. J. Wagner, 1910
San José for Helicina (Tristramia) punctisulcata cuericiensis Richling, 2004
Veracruz for Helicina elata Shuttleworth, 1852
Veracruz for Helicina merdigera L. Pfeiffer, 1855
Veraguas for Helicina funcki L. Pfeiffer, 1849
Yucatan for Helicina lirata L. Pfeiffer, 1847
Yucatan for Helicina tenuis L. Pfeiffer, 1849
Holotype USNM 536026, identified as Helicina terryae Rehder, 1940
