original description
Seninski, K. (1905). Novyya Dannyya o neogenovych' plastach' Yugozapadnogo Zakavkaz'ya. <em>Trudy Obshchestva Estestvoispytateley pri Imperatorskom Yuryevskom Universitete.</em> 16, 1-80.
page(s): 47-48, pl. 2, figs 28-31 [details]
From editor or global species database
Type locality "Pakveski" [P'ok'veshi] (Abkhazia) [details]